Alcidedorbignya inopinata (Muizon and Marshall 1992 Muizon et al 2015; MHNC 8372, Early Paleocene, 64 mya; scale bar is 2cm) was originally considered a basal pantodont (Condylartha) close to Pantolambda and the origin of ungulates and digging placentals. The rostrum is curved with a smaller naris. Long hooves are former claws on a semiplantigrade manus.

Metacheiromys tatusia (M. marshi (YPM 12903 holotype is chimaera, part primate; Middle Eocene; Wortman 1903, Osborn 1904, Simpson 1931, AMNH 11718, 25cm presacral length) was originally considered a primate, then an edentate and then as a palaenodont. Here the data nests between Prodinoceras and Ernanodon. Few to no teeth line the jaws, except for the upper and lower canines, which were retained. The sacrum appears odd in that only the posterior sacral ribs attached to the ilia. This may not be so.

Miacis parvivorus (aka Messelogale kessleri Cope 1872; Springhorn 1982, Springhorn 2000, Heinrich et al. 2008; 30cm in length; Late Paleocene-Late Eocene,) was considered a pre-carnivore, but here hests with Alcidedorbignya and Metacheiromys near the base of the pantodonts.

Ectoconus ditrigonus (Cope 1884 Early Paleocene, 66-63mya, sheep-sized) is tradiitionally considered a basal condylarth known from a rare complete skeleton. Here Ectoconus is derived from the marsupials, Alcidedorbignya and Monodelphis. It is a sister to Pantolambda. Though canines were present, Ectoconus was an herbivore with five digits on all four limbs. Shown here digitigrade, this taxon was likely plantigrade. Another species, Ectoconus majusculus, is shown here.