Glironia venusta (Thomas 1912; snout-vent length 20cm) is the extant bushy-tailed opossum. It is nocturnal, arboreal, small and omnivorous. Glironia lacks a marsupium (= pouch), by convergence with several marsupials, but basal to Placental 2 taxa with Ptilocercus at the base. The upper canine is a vestige. The first premolar has the appearance of a canine.

Dromiciops is another relative. Following tree shrews, placental moon rats, shrews, tenrecs, archaeocetes and odontocetes are descendant taxa in the LRT.