Nasua and Ambolestes

Nasua nasua (Linneaus 1766) is the extant coatimundi or coati. Here it nests with the Notharctus, as a basalmost primate. Traditionally nested with raccoons (Procyon) Nasua has three molars, all similar, and no carnasial tooth, distinct from all members of the Carnivora in the LRT.

Ambolestes

Ambolestes zhoui (Bi et al. 2018, Early Cretaceous, 126 mya; 25cm in length) is is one third the size of the extant opossum, Didelphis. In the LRT nests Ambolestes close to Eomaia, but clloser to Vincelestes.

Bi said in an interview, “Ambolestes zhoui is an early member of the placental lineage. It also carries mixed features both placentals and marsupials”. That is supported by the LRT.

Bi S-B, Zheng X-T, X, Wang X-L, Cignetti N-E, Yang SL and Wible JR 2018. An Early Cretaceous eutherian and the placental/marsupial dichotomy. Nature (advance online publication) DOI: https://doi.org/10.1038/s41586-018-0210-3
https://www.nature.com/articles/s41586-018-0210-3

Linneaus C von 1766. Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio duodecima, reformata. pp. 1–532. Holmiæ. (Salvius)

wiki/Nasua
wiki/Ambolestes

Mammalia - Monotremata

Ornithorhynchus

Marsupiala - Metatheria

Eomaia

Placentalia 1.1

Phenacodus

Placentalia 1.2

Carnivora

Nasua

Panthera

Smilodon