Sophineta cracoviensis (Evans and Borsuk-Bialynicka 2009) Early Triassic ~250 mya ~1 cm skull length was orginally considered a sister to living squamates (Iguania + Scleroglossa). Here Scandensia was derived from a sister to Paliguana and Santaisaurus at the base of the Lepidosauriformes.
Distinct from Paliguana, the skull of Sophineta was narrower overall and had a straight jawline. Sophineta had a taller naris and a longer postorbital region. TThe ascending process of the maxilla was taller. Teeth extended below the orbit. The posterior palate is preserved.
Sophineta is known from fissure fill fossils. Each bone is in 3D but the skeleton is completely disarticulated.
See the complete tree.