Protictitherium crassum (Kretzoi 1938; Koufos and Konidaris 2011; Miocene-Pliocene) is traditionally considered a hyena-like civet, but civets are not related to hyenas. Here Protictitherium nests with Protictis, basal to the anagalids. These two are among the first tree shrews to become terrestrial. |