Cryptolacerta hassiaca (Müller et al. 2011) Eocene, ~7 cm snout-to-vent length was orginally described as a precursor to burrowing amphisbaenians, like Bipes and Amphisbaena. Derived from a sister to Acanthodactylus and Chalcides, here Cryptolacerta nested at the base of the Heloderma clade, which also gave rise to several legless burrowing reptiles including Cylindrophis and Uropeltis.
The reconstruction of Müller et al. (2011, right above) could not be replicated. A DGS tracing (left above) produced a deeper skull with upper temporal fenestrae, an premaxillary ascending process and a lacrimal among several other differences.
Distinct from Chalcides, the skull of Cryptolacerta had a higher rostrum due to a higher maxilla, but Tupinambis and Acanthodactylus had a high maxilla. The postorbital was larger and more elongated. The posterior parietal was wider than the anterior. The premaxilla tucked under than appeared above the nasal. The jugal was gracile. The quadrate was straighter and taller.